Luperina nickerlii gueneei Doubleday, 1864

First discovered at Rhyl in 1860 or 1861
Habitat: sand dunes and beaches in north Wales and Lancashire
Foodplant: Elytrigia juncea (Sand Couch Grass)

	August 1998: adult gueneei hiding in Elytrigia clump by day
	August 1996: adult gueneei on Elytrigia at night
	26th August 2002: adult gueneei on sand
	20th March 2003: gueneei habitat, showing extensive dune system in north Wales